Category Archives: Articles

Two new species of genus Cousinia (Asteraceae, Cardueae) from W and NE Iran

Two new species of Cousinia are described from Iran. 
Cousinia bornmulleriana from the west that can be placed in sect. Haussknechtii based on some key characters such as flower numbers, phyllary numbers and the inflorescence form traits. This species can be also compared with its closest relative C. haussknechtii. and C. karkasensis.
The second new species, C. semnanensis, from North East of Iran, belongs to the sect. Stenocephalae for its flower numbers, winged stem, and phyllaries shape, and is compared to C. alexeenkoana and C. glaucopsis as its close relative.

To download the article please click here (DOI: 10.29252/nbr.9.2.144).

Cite as: Attar F, Maroofi H, Mirtadzadini M, Sotoodeh A. Two new species of genus Cousinia (Asteraceae, Cardueae) from W and NE Iran. nbr. 2022; 9 (2) :144-152


Verbascum birjandense and V. urumiense (Scrophulariaceae), two new endemic species for Flora of Iran

Abstract. Verbascum birjandense and V. urumiense are described as new endemic species of Verbascum from eastern and northwestern Iran. It was not possible to assign the samples of these two taxa to any of the known Verbascum species in Iran and adjacent regions. Collating the morphological characters of the samples against the specialized and pertinent literature of the genus led us to propose them as new species. Both belong to section Bothrosperma, subsection Fasciculata.
These new species are morphologically close to V. erianthum and V. songaricum based on some key characteristics, but show some differences in basal leaves shape, pedicel size, the ratio of pedicel to calyx, presence of hairs inside the corolla, and the hair color of the stamen’s filaments.

Cite as: Sotoodeh A., Attar F., Laitung B., Civeyrel L. (2022). Verbascum birjandense and V. urumiense (Scrophulariaceae), two new endemic species for Flora of Iran. Phytotaxa, 538: 035–044.

in ResearchGate

Potentilla alborzensis Faghir & Attar

A new species of the genus Potentilla L. (Rosaceae) from the Tehran province (Iran)

Potentilla alborzensis Faghir & Attar, sp. nov. (Rosaceae) is collected from steppe Mountains of Tang-e Washi, in the middle of Alborz Mountains for the first time. The new species is related to section Aureae (Rydb.) Juz. for its palmate radical leaves, having straight hairs and glandular trichomes; corymbiform inflorescence; hairy and glandular sepals; clavate and papillose style, shorter than or as long as matured ovary. Detailed micro- and macromorphological descriptions of the newly collected species, as well as photographs and distribution map are provided. In addition, comparison of diagnostic traits with allied species especially P. crantzii (Crantz) Beck ex Fritsch, P. adenophylla Boiss. & Hohenack., P. gelida C. A. Mey., P. geranioides Willd. and identification key are presented.


Onosma new species

Onosma khorramabadensis, O. lorestanica, O. sahandica and O. kerendica are described and photographed as new species from Lorestan, Eest Azarbayejan and Kermanshah provinces of Iran.

Cite as: Attar F., Sotoodeh A., Mirtadzadini M. (2020) Four new species in subsection Asterotricha for the genus Onosma L. (Boraginaceae) from flora of Iran. Feddes Repert, 131: 146-158. 

Some articles from 2018 and 2019


Hadidchi, A., Attar, F., & Ullah, F. (2020). Using microscopic techniques for taxonomic implications of seed and fruits of Delphinium L.(sensu lato)(Ranunculaceae). Microscopy Research and Technique83(2), 99-117.

Attar, F., Sotoodeh, A., & Mirtadzadini, M. (2019). Teucrium elymaiticum (Lamiaceae): a new species for Flora of Iran. Mediterranean Botany40(2), 203-207.

Moradi, H., & Attar, F. (2019). Comparative study of floristic diversity along altitude in the northern slope of the central Alborz Mountains, Iran. Biodiversitas Journal of Biological Diversity20(1), 305-312.

Attar, F., & Rad, M. A. (2019). Cousinia sharifii, a new species of Cousinia sect. Cynaroideae (Asteraceae) from NW Iran. Phytotaxa395(4), 287-295.

Niaki, N. A. R., Attar, F., Sheidai, M., Maroofi, H., Joharchi, M. R., & Ullah, F. (2019). Two new species of Cotoneaster (Rosaceae, subgen. Chaenopetalum) for the flora of Iran. Phytotaxa402(4), 207-215.

Attar, F., Esfandani‐Bozchaloyi, S., Mirtadzadini, M., Ullah, F., & Zaman, W. (2019). Foliar and stem epidermal anatomy of the tribe Cynoglosseae (Boraginaceae) and their taxonomic significance. Microscopy Research and Technique82(6), 786-802.

Niaki, N. A. R., Attar, F., Mirtadzadini, M., Mahdigholi, K., & Sheidai, M. (2019). Micromorphological studies of the leaf epidermis of Iranian Cotoneaster Medik. NORDIC JOURNAL OF BOTANY37(2).


Attar, F., Mozaffarian, V., & Mirtadzadini, M. (2018). Cousinia elymaitica (Cardueae, Asteraceae) a new species from west of Iran. Nova Biologica Reperta5(3), 307-310.

Rastegar, A., Attar, F., & Mirtadzadini, M. (2018). Novelties in Iranian Cousinia sect. Cynaroideae (Asteraceae, Cardueae): new taxa and taxonomic notes. Phytotaxa343(3), 227-239.

Attar, F., Esfandani-Bozchaloyi, S., Mirtadzadini, M., & Ullah, F. (2018). Taxonomic identification in the tribe Cynoglosseae (Boraginaceae) using palynological characteristics. Flora249, 97-110.

Sotoodeh A., Attar F.,Andalo C., Mirtajaldini M. And L. Civeyrel. (2018). Focusing on three Verbascum L. taxa (Scrophulariaceae) of the Flora of Iran. Adansonia, 40:171-181.

Focusing on three Verbascum L. taxa (Scrophulariaceae) of the Flora of Iran

Verbascum carmanicum (Bornm.) Hub.-Mor. subsp. carmanicum
Verbascum carmanicum (Bornm.) Hub.-Mor. subsp. kermanense (Hub.-Mor.) Sotoodeh, Attar, Andalo & Civeyrel, comb. nov., stat. nov.
Verbascum gabrielae (Bornm.) Hub.-Mor.

Focusing on three Verbascum L. taxa (Scrophulariaceae) of the Flora of Iran
Sotoodeh A., Attar F., Andalo C., Mirtadzadini M. & Civeyrel L. 2018. — Focus sur trois taxa de du genre Verbascum L. (Scrophulariaceae) de la Flore d’Iran. Adansonia 40 (13): 171-181.